I have seen the same thing in our axolotls when we have attempted to
chill them and stop development for videp taping so I suspect your
problem is due to messing up some aspect of their development rather than
an effect of your operation. We stopped dropping the axolotls below 8
degrees and the problem vanished.This does mean we need to work faster
because they do keep developing. I would be very interested in knowing if
anyone has thoughts on why this occurs.
Natalie K Bjorklund