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BEN # 266

Adolf Ceska aceska at victoria.tc.ca
Tue Mar 27 02:56:45 EST 2001

BBBBB    EEEEEE   NN   N             ISSN 1188-603X
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BBBBB    EEEEE    NN N N             BOTANICAL
BB   B   EE       NN  NN             ELECTRONIC
BBBBB    EEEEEE   NN   N             NEWS

No. 266                              March 26, 2000

aceska at victoria.tc.ca                Victoria, B.C.
 Dr. A. Ceska, P.O.Box 8546, Victoria, B.C. Canada V8W 3S2


The  Central  Alaska  network  of  the  National Park Service is
currently seeking applications for a Botanist position to assist
in leading a vascular plant inventory study in three of Alaska's
premier National Park units: Denali National Park,  Wrangell-St.
Elias National Park and Yukon-Charley Rivers National Preserve.

The person in this position will take part in all aspects of the
inventory  study  including  site selection, fieldwork, specimen
identification and data analysis.  This  is  truly  an  exciting
opportunity  for  field-oriented botanists. This job opportunity
is posted on the USA jobs website of the United States Office of
Personnel Management  (http://www.usajobs.opm.gov).  Application
instructions  and  a  more  complete description may be found by
performing an "agency job  search"  for  Alaska  Region  of  the
National Park Service at this web address!!

For  information about this opportunity, please call the Susanne
Brown, personnel specialist for Denali National  Park  at  (907)
683-9503.  You  may  also  contact  the project principal inves-
tigator, Carl Roland at (907) 456-0479.

From: Dr. Bill Weber [weberw at colorado.edu]

Leon Kelso was a government employee in Washington, D.C. at  the
Bureau  of Biological Survey. His specialty was ornithology, but
he was also very much interested in the Rocky Mountain Flora. He
never published in a "refereed journal," and  was  probably  the
last  of  a breed of American scientists who felt he had to pub-
lish his own journal. This was a series which he called Biologi-
cal Leaflets. It appears that he published at least 88 of these,
on topics ranging from the taxonomy  of  owls,  the  anatomy  of
feathers,  the  Colorado flora, the willows (_Salix_) and sedges
(_Carex_) in Colorado.

Very few sets of the Biological Leaflets remain. The  University
of  Colorado  Herbarium in Boulder has had some of them for many
years, but I recently discovered many missing numbers among  the
papers  of  Askell Love, who must have carried on correspondence
with Kelso over many years.

I am lacking the following numbers:

2, 3, 4, 7, 8, 12, 13, 22, 27, 46, 72, 73, 75, 76, 77,  80,  81-
85, 87.

I  am  putting together a paper about him and would like to have
photocopies of the missing papers, which are often only  a  page
or two long. Probably these are not all botanical papers, for he
published  a lot about birds, feathers, and bioelectronic obser-
vations having to do with the migrations, homing, etc. of birds.

Thanks for that!


Dr. W.A. Weber
University of Colorado Museum
Campus Box 315
Boulder CO 80309  USA

From: Stephen Timme [slt at pittstate.edu]

Malcolm,  Bill  &  Nancy  Malcolm.  2000.   Mosses   and   other
   bryophytes:  An  illustrated  glossary.  Micro-Optics  Press,
   Nelson, New Zealand. iv + 220  p.  ISBN  0-473-06730-7  [hard
   cover] Price: US$39.95

   Published by:
   Micro-Optics Press,
   a division of Micro-Optics Ltd.
   Box 320, Nelson, New Zealand
   Available from:
   Timber  Press,  Inc.  The Haseltine Building,
   133 S.W. Second Ave., Suite 450,
   Portland, OR 97204, USA
   Tel.: 1-800-327-5680 or 1-503-227-2878,
   Fax: 1-503-227-3070
   Web: http://www.timberpress.com

The Malcolms have produced a glossary of bryophyte terms that is
long  overdue.  This  illustrated  glossary  is  packed with the
majority of terms associated with moss, liverwort  and  hornwort
characteristics.  Most  are  illustrated  with color plates. For
characteristics that are highly variable,  the  authors  include
several  color  plates  to  illustrate the variability among the
bryophytes. These should give at least the beginning student  of
bryology  a better feel for the characteristic under study. Each
plate is titled by species name, characteristic (in  bold-face),
and  a  scale  of measurement. There is a brief introduction and
following the text a reference to some  literature  for  further
reading.  This  is  followed by an index to the illustrations by
species name. The results of this text is obviously the  efforts
of  years  of study and developing photographic techniques. This
reference will ease the frustrations often  encountered  by  the
beginning  student  of  bryology.  It  will also be an excellent
reference to the trained bryologist as well.

[This book contains 970 full-colour  close-ups  and  micrographs
plus  22 drawings illustrating 1550 cross-referenced entries and
400 species, a  comprehensive  glossary  and  a  celebration  of
diversity of bryophytes.]

From: Adolf Ceska [aceska at victoria.tc.ca]

Brayshaw, T.C. 2000. Pondweeds, bur-reeds and their relatives of
   British  Columbia:  Aquatic  families of monocotyledons. [2nd
   edition] Royal British  Columbia  Museum,  Victoria.  250  p.
   ISBN 0-7718-9574-7 Price: CDN$ 24.95

   Available from your local book stores. 
   If not, they can order it for you from:
   Raincoast Books
   9050 Shaughnessy Street
   Vancouver B.C., CANADA V6P 6E5
   Phone: 604 323-7100, Fax: 604 323-2600
   E-mail: info at raincoast.com

This  is  a  revised  edition  of the 1985 publication (Brayshaw
1985) that was originally  published  by  the  British  Columbia
Provincial  Museum  as  Occasional  Paper Series no. 26. It is a
treatment of 14 families of monocots with most of their  members
being  aquatic.  In  this  edition,  Dr.  Chris Brayshaw deleted
several species erroneously reported in the first edition (e.g.,
_Potamogeton diversifolius_, _Sparganium glomeratum_), and added
several species that were not known in British Columbia when the
first edition was published  (e.g.,  _Elodea  callitrichoides_).
The  discussion of many species has been expanded and the author
added new taxonomical treatments of  certain  species,  such  as
_Potamogeton filiformis_ and _Triglochin maritima_.

Chris Brayshaw's book appeared almost simultaneously with Crow &
Hellquist's  Aquatic  and  wetland  plants of northeastern North
America (Crow & Hellquist 2000), Flora of North America Vol.  22
(FNA  editors 2000), and Kartesz' Synthesis (Kartesz 1999). When
I tried to review Chris Brayshaw's book, I found myself  review-
ing these three publications and a CD-ROM, all at once.

All works more or less agree with each other in the treatment of
the  genus _Potamogeton_, although Brayshaw was the only one who
did  not  accept  the  separation  of   _Potamogeton_   subgenus
_Coleogeton_  in  the genus _Stuckenia_. I am a strong proponent
of using the stem anatomical characters  for  identification  of
broad-leaved  pondweeds,  but  none of the recent North American
treatments (except Ceska 1994 and 2001) employs  this  technique
as  a  routine identification tool. Brayshaw is the only one who
mentions its usefulness for identification of "eroded or  other-
wise  mutilated  material."  I  believe that in British Columbia
_Potamogeton nodosus_ is less common than shown on the distribu-
tion map. It is restricted to  the  lower  Fraser  River  Valley
(Hatzic  Lake)  and  the  Okanagan  Valley and plants from other
areas may belong either to another species or to _Potamogeton  x
sparganiifolius_  =  _P.  natans  x P. gramineus_. Morphological
characters of this complex are sometimes so subtle that one  has
to  use stem anatomical characters in order to reliably identify
even the best developed specimens  (Ogden  1943,  Wiegleb  1990,
Ceska  1994,  2001).  I have seen many specimens of broad-leaved
pondweeds that were annotated by E.C. Ogden (who developed  this
identification  technique)  and  most  of  them had a small pre-
printed slip on which Ogden  marked  his  observations  of  stem
anatomical characters.

The  genus  _Stuckenia_  (=_Potamogeton_  subgen.  _Coleogeton_)
requires major revision  worldwide.  With  the  introduction  of
_Stuckenia  filiformis_ subsp. _occidentalis_ (Chris Brayshaw is
possibly correct when he considers this subspecies a  hybrid  of
_S.  filiformis  x  S.  vaginata_),  many  specimens  previously
treated as _S. vaginata_ would belong to  this  subspecies.  The
boundary  between  _S. filiformis_ subsp. _occidentalis_ and _S.
vaginata_ is blurred and quite subjective. We need more  studies
of  this  complex,  and  the  further  study  may  show that _S.
vaginata_ is much less common in British Columbia than shown  on
Chris Brayshaw's map.

In  the  Flora  of  North  America treatment, Haynes & Hellquist
reduced _Triglochin concinna_ to  small  plants  of  _Triglochin
maritima_.  I  am  sure that botanists from the Pacific Coast of
North America will consider this a gross mistake. In his  _Trig-
lochin_  treatment,  Chris  Brayshaw  retains _T. concinna_. The
question is what _Triglochin debilis_ is. Chris Brayshaw  inves-
tigated  the  type of this taxon and concluded that _T. debilis_
can be regarded a variety of  _T.  concinna_.  Brayshaw  devotes
about  two  pages  to  the  analysis  of  the  specimen  of "_T.
debilis_" (V 170,988) that  I  and  my  wife  collected  at  the
Osoyoos  Lake. I was attracted to this plant when I saw Bill van
Dieren's collection of similar  plants  from  the  Shuswap  Lake
area. I concur with Chris Brayshaw that more collecting and more
taxonomic  work  in  _Triglochin  maritima - concinna - debilis_
complex should be done. Kartesz (1999) recognized _T.  concinna_
as  a  species  distinct  from  _T.  maritima_,  but treated _T.
debilis_ as a mere synonym of _T. concinna_. I agree that it  is
necessary  to  re-evaluate "_Triglochin debilis_" as well as the
anomalous specimens from the Interior British Columbia.

In his 1985 treatment of _Sparganium_, Chris Brayshaw  published
three new combinations in _Sparganium angustifolium_. He treated
_Sparganium    emersum_   as   a   subspecies   of   _Sparganium
angustifolium_. In turn, he treated  _Sparganium  chloropetalum_
as  a  variety  of  _S. angustifolium_ subsp. _emersum_, and _S.
multipedunculatum_ as a variety  of  _S.  angustifolium_  subsp.
_angustifolium_.  Cook  & Nicholls (1986) recognized _Sparganium
angustifolium_ and _S. emersum_  as  two  distinct  species  and
synonymized  _S.  multipedunculatum_ with _S. angustifolium_ and
_S. chlorocarpum_ with  _S.  emersum_.  Whilst  Flora  of  North
America  and  Crow  & Hellquist (2000) followed Cook & Nicholls'
treatment, Brayshaw (2000) follows his original  1985  treatment
of  a  single  species  with  two subspecies and four varieties.
Kartesz (1999) reduced all four taxa into _S. angustifolium_ and
listed  Chris  Brayshaw's  subspecies  and  varieties  as   mere
synonyms. Consequently, _Sparganium emersum_ got completely lost
from  North  America  in  Kartesz'  "Synthesis".  In my opinion,
_Sparganium angustifolium_ and _S.  emersum_  are  two  distinct
species  and  it  was a mistake that Brayshaw (2000) and Kartesz
(1999) did not follow the _Sparganium_  treatment by  Cook   and
Nicholls (1986).

Chris  Brayshaw has an enviable gift of being able to do his own
botanical illustrations. It is a great asset when a botanist can
transform his ideas into precise  botanical  illustrations,  and
Chris Brayshaw's illustrations belong to the best in its class.

The  Royal  British Columbia Museum has to be commended for pub-
lishing this treatment. I liked the return to the  format  close
to  the  B.C.  Provincial  Museum's  favourite "Handbook" format
(although these new "Handbooks" are few inches larger  than  the
original  Handbooks  and  the  assignment to any series has been
dropped). It is a pity that the manuscript was  not  more  care-
fully  edited  and  reviewed  by  a  plant  taxonomist  to avoid
misspellings  such  as  "stiff-leafed"   and   other   "-leafed"
pondweeds  (I  have  never  dreamt about having to correct Chris
Brayshaw's English!), citing  _Triglochin  debilis_  (M.E.Jones)
Love  and  Love as a synonym of one species and its basionym _T.
maritima_ var. _debilis_ M.E. Jones  as  a  synonym  of  another
species,  etc.  In spite of these shortcomings, Chris Brayshaw's
book is a significant contribution to the knowledge  of  aquatic
monocots in British Columbia and the Pacific Northwest.


Brayshaw,  T.C.  1985.  Pondweeds  and bur-reed, and their rela-
   tives: Aquatic families of monocotyledons in  British  Colum-
   bia.  Occasional  Papers  of  the British Columbia Provincial
   Museum, no. 26. Province of British Columbia, Victoria,  B.C.
   166 p.
Ceska,  A. 1994. Potamogetonaceae. Pp. 146-151, 177-178, 184 in:
   Douglas, G.W., G.B.  Straley  and  D.  Meidinger  [eds.]  The
   vascular plants of British Columbia. Part 4 - Monocotyledons.
   Special  Report  Series 2, B.C. Ministry of Forests, Research
   Branch, Victoria. 257 p. [See BEN # 75]
Ceska, A. 2001 [in press]. Potamogetonaceae. In: Douglas,  G.W.,
   J.  Pojar & D. Meidinger. Illustrated Flora of British Colum-
   bia. Vol. 7.
Cook, C.D.K. & M.S. Nicholls. 1986. A monographic study  of  the
   genus  Sparganium  (Sparganiaceae).  Part  1.  Subgenus  Xan-
   thosparganium Holmberg. Bot. Helvetica 96: 213-267.
Crow, Garrett E. & C. Barre Hellquist. 2000. Aquatic and wetland
   plants of northeastern North America: a revised and  enlarged
   edition  of  Norman  C. Fassett's A manual of aquatic plants.
   Volume 2:  Angiosperms:  monocotyledons.  The  University  of
   Wisconsin Press, Madison. LV+400 p. [See BEN # 261]
Flora  of  North  America,  Editorial  Committee. 2000. Flora of
   North America North  of  Mexico,  Volume  22,  Magnoliophyta:
   Alismatidae,   Arecidae,  Commelinidae  (in  part)  and  Zin-
   giberidae. Missouri Botanical Garden Press,  St.  Louis,  MO.
   xxii + 352 p.
Kartesz,  J.T.  &  C.A.  Meacham.  1999.  Synthesis of the North
   American Flora, Version 1.0. North Carolina Botanical Garden,
   Chapel Hill, NC. [CD-ROM] [See BEN # 237]
Ogden, E.C. 1943. The broad-leaved  species  of  Potamogeton  of
   North  America  north of Mexico. Rhodora 45: 57-105, 119-163,
Wiegleb, G. 1990. The importance of stem  anatomical  characters
   for  the  systematics  of the genus Potamogeton L. Flora 184:

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