please find enclosed the final program of the EUROSILVA Workshop on
'Development and ageing in forest trees', to be held in Florence, Italy,
on 20-24 September, 2000.
The program can also be accessed on the workshop web page at:
Please circulate it as widely as possible.
Prof. Marco Borghetti
Prof Giuseppe Scarascia Mugnozza
Dr Federico Magnani
Workshop local organisers
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DEVELOPMENT AND AGEING
IN FOREST TREES
20-24 September, 2000
Please visit the Workshop web page at:
What controls tree development and ageing? what is the relative role of internal and external factors? At the tissue level, recent advances have demonstrated in detail that development and senescence both result from the interaction of genetic and environmental controls, involving differential gene expression and activation. At the tree level, a shift in hormonal patterns, possibly of endogenous genetic origin, has been found to trigger the process of maturation, which strongly affects the structure, function and growth of the plant and initiates the diversion of resources from vegetative to reproductive growth. Also at the tree and population level, exogenous factors have been proposed to control the developmental pattern of rise and decline of primary productivity with age. A lively discussion has been going on for several years on what could be the ecological determinants of the age-related decline in forest productivity. Several theories have been proposed to explain this phenomenon, variously suggesting nutrient or hydraulic limitations, increased respiration and shifts in biomass allocation as possible mechanisms, but to date there is no definitive evidence to support any of them as a universal mechanism. Ageing and maturation are generally thought to be intimately interwined albeith distinct processes. Eventually, what are the implications of development and ageing for the response of tissues and plants to their biotic and abiotic environment?
Because of the variety of processes involved in tree development and maturation, definite answers often result elusive, requiring a comprehensive and holistic approach that takes both endogenous and external factors into account.
The aim of the workshop is therefore to bring together experts of different background and expertise to discuss these processes of central relevance both to plant biology and to forest ecology moving from different perspectives. An interdisciplinary workshop on this issue that brings together foresters, ecologists and biologists seems most appropriate and relevant to the aims of the EUROSILVA Action.
TREE PHYSIOLOGY SPECIAL ISSUE
Tree Physiology, a well-known refereed journal, has agreed to draw from the Workshop a special issue on 'Development and ageing of forest trees'. All the authors of accepted oral presentations are invited to contribute, although only those papers that are deemed acceptable by the reviewers and editor will be published in the special issue of Tree Physiology.
Papers should be prepared in accordance with Tree Physiology's guide for contributors and submitted (original plus three copies) to the workshop local organisers no later than 31 October, 2000. The length of the manuscript should not exceed 8 pages (about 8000 words, including references). The manuscript should also be provided as a digital file (a PDF file, if possible, otherwise an MS Word or Word Perfect file, on an IBM-format floppy disk, zip disk or CD-ROM). The file should be named after the corresponding author, e.g. smith.doc, and contain all components of the article including tables, figures and figure captions.
WORKSHOP FINAL PROGRAMME
Wednesday, 20 September
15.00 Arrival and registration
17.30 Opening session
18.30 Working group meeting
Thursday, 21 September - WG1 Growth and development
8.30 Valjakka M., Tuhkanen E., Vapaavuori E., H?ggman H., Kangasjarvi J.
Keynote: Gene expression during leaf development and senescence in birch (Betula pendula)
9.00 Rohde A., Boerjan W.
Keynote: Molecular components of terminal bud formation in poplar
9.30 Palva E.T., Heino P., Li C.Y., Puhakainen T., Welling A., Boije M., Aalto O.
Development of dormancy and winter hardiess in birch
9.50 Collins A., Jones H., Campbell M.
The control of meristem establishment, maintenance and maturation in Eucalyptus
10.40 Moritz T., Eriksson M., Israelsson M., Sandberg G., Olsson O., Olsen J., Junttila O.
Keynote: Studies of photoperiodic induction of shoot elongation in trees: a transgenic approach
11.10 Magel E., Sundberg B., Uggla C.
Sucrose synthase is the dominating sucrose cleaving enzyme in secondary differentiation processes of forest tree axes, cambial growth and heartwood formation
11.30 Piispanen R., Saranpaa P.
Seasonal variation in storage lipids and non-structural carbohydrates of young silver birch (Betula pendula Roth) trees
11.50 Taylor G., Robinson K.M., Ferris R., Bunn S.M., Bradshaw H.D.
Leaf growth and yield in poplar: can putative QTL be identified?
14.00 Poster session
15.00 Drouet A.
Keynote: Ageing control on flavonoid biosynthesis in walnut
15.30 Diaz-Sala C.
Keynote: Maturation and rejuvenation in trees: physiological, molecular and cellular mechanisms
16.00 Barnett J.R., Eshtiaghi Z.
Morphological and anatomical rejuvenation induced by sequential grafting in walnut
16.50 Fraga M.F., Centeno M.L., Ca?al M.J., Rodr?guez A., Fern?ndez B., Rodriguez R.
Molecular basis of organ and tree ageing and reinvigoration: applications to micropropagation of selected progenies and mature trees of Pinus radiata D. Don
17.10 Ritter E., Fernandez B., Rodriguez R., Giannino D., Racchi M., Wagner E., Albrechtowa J., Favre J.M., Smulders S.M.J., van der Linden G., Pasqualetto P.L., Paques M.
Development, validation and application of molecular, morphological and physiological markers for juvenile and mature state characterisation in woody plant species (FAIR3- CT96-1445)
17.30 Danti S., Bagnoli F., Caparrini S., Racchi M.L.
Catalases are differentially expressed during development in peach (Prunus persica)
17.50 Poster session
18.30 MC meeting
Friday, 22 September - WG2 Mineral nutrition and water relations
8.30 Mencuccini M.
Keynote: Functional interpretation of allometric analyses in forest trees. The role of hydraulic constraints
9.10 Kostner B., Falge E., Bernhofer Ch., Tenhunen J.
Age- and management-related effects on leaf area / sapwood area relationships, canopy transpiration, and carbon gain of Picea abies stands in Central Germany
9.30 Magnani F., Mencuccini M., Borghetti M.
Optimal self-similarity in xylem structure. Implications for the developmental pattern of tree hydraulic architecture and gas exchange
9.50 Hubbard R.M., Stiller V., Ryan M.G., Sperry J.S.
Stomatal conductance and photosynthesis vary linearly with plant hydraulic conductance in ponderosa pine
10.40 Gower S.T.
Keynote: Towards a better understanding of age-related forest NPP decline
11.20 Bond B.J., Ryan M.G., Phillips N., McDowell N.G.
Testing the hydraulic limitation hypothesis in tropical and temperate ecosystems
11.40 Nikinmaa E., M?kel? A., Berninger F., Hari P., Peram?ki M., Valentine H.
Size related decline of tree productivity from carbon balance perspective
14.00 Poster session
14.40 Ryan M.G., Bond B.J., Hubbard R.M., Williams M.
Keynote: Experimental evidence for hydraulic constraints on stomatal function, and implications for age-related decline in forest growth
15.20 Rust S., Roloff A.
Reduced photosynthesis in old oak (Quercus robur L.): the impact of crown and hydraulic architecture
15.40 Niinemets U.
Changes in foliar morphology and chemical composition with increasing tree size: a review of several case studies
16.00 Giardina C., Ryan M., Fownes J., Binkley D.
Total belowground carbon allocation and aboveground net primary production in a fast growing Eucalyptus plantation
16.50 Urbinati C., Carrer M., Anfodillo T.
Age-dependent radial growth differences of Norway spruce (Picea abies Karst.) in Eastern Italian Alps
17.10 Vanninen P.
Carbon budget for individual Scots pine trees: effects of size, competition and site fertility on growth allocation
17.30 Bauer G.A., Bazzaz F.A.
Nitrogen partitioning influences photosynthetic capacity during leaf aging
17.50 Poster session
18.30 MC meeting
Saturday, 23 September - WG3 Biotic and abiotic interactions
8.30 Ceulemans R., Bortier K., Gielen B., Laureysens I., Janssens I.A.
Keynote: Developmental processes in trees: effects of abiotic factors, competition and genotype
9.10 Marek M.V., Sprtova. M.
Thinning effects on photosynthetic characteristic of a Norway spruce stand
9.30 Stewart J.
Understanding the biological processes underlying growth stagnation in lodgepole pine and its response to silvicultural interventions
9.50 Wirth C., Schulze E.-D.
Multiple control of variability of above-ground NPP in Siberian Scots pine forests
10.40 Kuppers M.
Keynote: Ecophysiology of forest succession: from leaf to plant in a competitive environment
11.20 Valentini R., Manca G., Dore S., Tedeschi V., Tirone G.
Age related carbon dynamics in a Quercus cerris L. coppice forest under intensive management
11.40 Carrer M., Urbinati C., Anfodillo T.
Age influence in tree ring growth response to climate of Larix decidua Mill. and Pinus cembra L. at timberline
14.00 Poster session
14.40 Kolb T.
Keynote: Ageing as an influence on tree response to ozone. Theory and observations
15.20 Wieser G.
Can tree age dependent differences in ozone susceptibility of Picea abies be related to cumulative ozone uptake and ozone flux ?
15.40 Havranek W.M., Wieser G., Tegischer K.
Ozone effects in a multiple stress experiment with young Norway spruce
16.00 Vapaavuori E., Riikonen J., Oksanen E., Repo T., Peltonen P., Holopainen T., Holopainen J., Julkunen-Tiitto R.
Responses of fast-growing ozone-tolerant and ozone-sensitive silver birch clones to elevated CO2 and O3: clonal differences and results from first year of fumigation
16.50 Scarascia Mugnozza G., Calfapietra C., Sabatti M., de Angelis P., Ceulemans R., Gielen B., Miglietta F.
Biomass growth and canopy development under elevated CO2 conditions of a poplar tree plantation: the POPFACE experiment
17.10 Ferris R., Sabatti M., Miglietta F., Mills R.F., Taylor G.
Leaf cell expansion and leaf cell production in poplar are both increased in elevated CO2: a free-air CO2 enrichment (POPFACE) study
17.30 Schutzendubel A., Langenfeld-Heyser R., Fritz E., Teichmann T., Ott, Godbold D., Polle A.
Cadmium-induced oxidative stress in ecomycorrhizal fungi and hosts
17.50 Poster session
18.30 MC meeting
Sunday, 24 September
9.00 Field excursion: Tuscanian forests and vineyards
18.00 Return to Florence
21 September - WG1 Growth and development
Abe H., Nakai T. Effects of the temporal deficit on morphogenesis of the xylem cells in Cryptomeria japonica D. Don
Benderoth G., Silber G., Koloupaev V. Mechanical limits for the growth of tree stems
Climent J., Chambel M.R., P?rez E., Gil L. Prediction of heartwood radius in Pinus canariensis Chr. Sm. Ex DC.
Funada R., Iwatate-Suzuki T., Utsumi Y., Suzuki T., Sano Y. Cambial reactivation in the
ring-porous hardwood Fraxinus mandshurica var. japonica
Giovannelli A., Giannini R. In vitro growth characteristics of mature and re-invigorated chestnut (Castanea sativa Mill.)
Palumbo M.G., Giannini R., Paffetti D. Characterization of different growth stages in Cupressus sempervirens L.
Kauppinen L., Immanen J., Ulvila J., Paulin L., Palva T., Helariutta Y. Analysis of wood
development in birch by cDNA sequencing
Kurth W., Anzola J?rgenson G., Dzierzon H., Schulte M. Digital reconstruction of tree architecture as a tool for assessing growth and functional performance
Pirttila A.M., Laukkanen H., Hohtola A. Differences in the regeneration capacity of mature and juvenile Scots pine tissues
Saranpaa P., Piispanen R. Neutral and phospholipids in sapwood and heartwood of Scots pine (Pinus sylvestris L.)
Schaber J., Badeck F. Physiology-based phenology models of forest trees
Tegischer K., Tausz M., Grill D., Wieser G. Tree-age and needle-age dependent variations of antioxidants and photoprotective pigments in spruce needles
Valjakka M., Luomala E.-M., Vapaavuori E., Sutinen S., Kangasjarvi J., and Haggman H.
Photosynthesis, growth and senescence in sense-RbcS transformed birch (Betula pendula) lines
Zaspel I., Hertel H. Development of progenies of Quercus petraea and Q. robur descending from selected old trees of a relic population
22 September - WG2 Mineral nutrition and water relations
Anttonen S., Jolkkonen A., Linder S., Lundmark T., Vapaavuori E. Effects of optimal fertilisation on carbon allocation, and chemical composition of Norway spruce
Aranda X., De Herralde F., Fleck I., Sav? R. Hydraulic conductivity in Quercus ilex resprouts after fire
Gall R., Landolt W., Bucher J.B. Are reversible changes in bark size driven by the descent of assimilation products in Norway spruce stems?
Genenger M., Brodbeck S., Zimmermann S., Frossard E., Brunner I. Nitrate reductase activity of Norway spruce fine roots as affectedby nitrogen and wood-ash fertilisation
Iivonen S., Rikala R., Vapaavuori E. Seasonal patterns of root and shoot growth, gas exchange and carbohydrate status of Scots pine seedlings subjected to low or high nutrient availability
Kutnar L., Levanic T. Growth and age characteristics of the Norway spruce (Picea abies (L.) Karst.) from the mires on the Pokljuka Plateau, Slovenia
Landolt W., Michellod V., Bleuler P., Bucher J. Effects of soil fertility, elevated CO2 and increased N deposition on biochemical parameters in the foliage of young beech and spruce trees
Mencuccini M., Grace J. Age-related dynamics of carbon exchange in European forests: the EU Carbo-Age Project
Oberhuber W. Vulnerability of an inner Alpine drought-exposed forest ecosystem to climatic extremes
Patz G. The water balance in trees
Patakas A., Noitsakis B., Radoglou K., Jarvis P. Relationships between photosynthetic rate and leaf anatomy in two evergreen oak species
Radoglou K., Raftoyannis Y. Seasonal variation in physiological parameters of broadleaved
seedlings during the first two years of field establishment
Whitehead D., Ryan M.G. Low leaf-specific hydraulic conductivity reduces stomatal conductance and photosynthesis in older, taller mountain beech trees
Zitnik S., Muller C., Cl?Cment A., Bonnet-Masimbert M., Hanke D.E., Kraigher H. Metabolism of sugars and phytic acid during long term storage of acorns at low temperatures
23 September - WG3 Biotic and abiotic interactions
Einhorn K.S. Growth of beech (Fagus sylvatica) seedlings in response to temperature at four different light levels in simulated canopy gaps
Gielen B., Calfapietra C., Ceulemans R. Effects of elevated CO2 on crown structure, leaf area and growth of poplar genotypes in the POPFACE experiment
Kozovits A.R., Grams T.E.E., Blaschke H., Sommerkorn M., Matyssek R. Competition between
beech (Fagus sylvatica) and spruce (Picea abies) saplings under CO2/O3-regimes
Muller M., Stabentheiner E., Tausz M., Wonisch A., Grill D. Structural and physiological responses of forest tree species to different environmental situations
Reiter I., Haberle K.-H., Blaschke H., Matyssek R. Quantifying competition on crown level
between mature European beech and Norway spruce
Jokela A., Cordeiro A., Altabella T., Sarjala T., Bortolotti C., Tiburcio A., Huttunen S. Cloning and molecular analysis of arginine decarboxylase (ADC) from Scots pine
La Mantia T., Cullotta S., Marchetti M., Barbera G. Ecophysiology of holm oak (Quercus ilex L.) in different environmental conditions in Sicily
Lomas C., Stober C., George E. Chronosequence studies of tree fine root growth and turnover
Michelozzi M., Tognetti R., Rossi F. Seasonal variations of photosynthetic capacities and total chlorophyll contents in Aleppo pine trees from Italian provenances
Paoletti E., Raddi P., Di Lonardo V. Evolution of the Cupressus sempervirens - Seiridium
cardinale interaction in elevated CO2 over time
Petkovsek S.A.S., Kraigher H. Impact of pollution on biodiversity of types of ectomycorrhizae
Trost T., Gaberscik A. Responses of Norway spruce (Picea abies (L.) Karst.) to enhanced UV-B radiation
Vilhar U., Rupel M., Diaci J., Kraigher H. Norway spruce regeneration and interactions in the mycorrhizosphere
Please note: all forms and relevant information can be found on the Workshop web page at:
Workshop Venue and Secretariat
All the working sessions will be held at the Educatorio di Fuligno - Via Faenza, 48 - 50123 Florence (Italy) - ph. ++39 055 210232, a former convent newly transformed into conference venue, within walking distance of Florence city centre. The Secretariat will open at 2:30 p.m. on Wednesday, 20 September and will remain open for the duration of the Workshop. Accomodation will be in nearby hotels. Florence, possibly the most charming of all cities, can be easily reached either by plane or by train.
How to register
All participants must register by filling in the registration form (available on the workshop web page) and returning it, with payment, to the Organizing Secretariat Enic.
Registration Fees (VAT included)
By bank check/money order:
until 30 April 2000 - EUR 200
after 30 April 2000 - EUR 250
By credit card:
until 30 April 2000 - EUR 207
after 30 April 2000 - EUR 259
The fee includes: abstract book, congress kit, coffee breaks, working lunches and the social dinner. Please remember that registration forms received without payment will not be honored.
You may pay your congress fee by:
a) Bank check - payable to Enic/EUROSILVA
b) Money order - payment must be received by ENIC/EUROSILVA - account number 23884/00 (ABI/CAB 06160/02805), Cassa di Risparmio di Firenze, Agenzia n. 5, Via Gioberti n. 163/r, Florence, Italy. Payments are to be made net of all bank charges which are to be paid by the participant.
Note If you pay by bank transfer, please attach a copy of the transfer order to your registration form.
c) Credit card - we accept VISA, CARTA SI, MASTERCARD and EUROCARD. Please remember to fill in the pertinent sections of the registration form.
Refunds will be made after the closing of the workshop (upon written request submitted to the Organizing Secretariat Enic via fax) as follows:
before 30 June, 2000: 25% of the total amount will be withheld
after 30 June, 2000: no refunds will be made
A field excursion will be organized on Sunday 24 September, 2000. Local foresters and ecologists will take us through the Chianti region to the oak and beech stands of Montagnola Senese and of the Parco Naturale della Val d'Orcia, close to the Monte Amiata massif. We will then move on to visit the medieval abbey of Sant'Antimo, founded by Charlemagne in 871, and the medieval village of Montalcino, home to one of the most famous Italian wines, the Brunello. A guided tour of the cellars in one of the most interesting wineries in the area, complete with a wine tasting experience, will end the day. Please note that an additional EUR 50 will be charged for the field excursion.
The Organizing Secretariat Enic has reserved a sufficient number of rooms in hotels located near the Workshop venue. Please fill in the appropriate spaces on the registration form and return it to Enic. Reservations cannot be accepted without a first night deposit + EUR 13 for booking fees. The deposit, minus the booking fees, will be deducted from your hotel bill upon presentation of the voucher you will receive from Enic. Please note that the hotel balance must be paid in Italian Liras directly to the hotel upon departure.
Please specify your accomodation requirements in the registration form (available in Acrobat .pdf or MS-Word .zip format) and return it, along with the first night deposit for the category of your choice, no later than June 30th. We cannot guarantee room availability after that date since the number of rooms in downtown Florence is limited.
Rooms will be assigned on a first-come-first-served basis. If the single rooms are sold out, we will assign double rooms for single occupancy; when rooms are no longer available in the category of your choice, we will automatically book you into the next highest category. Your consent will be asked before taking any actions.
Cancellations: written cancellations received before July 30thwill be entitled to a refund of the first night deposit less EUR 26 for administrative fees. No refunds will be made for cancellations received after June 30th. All refunds will be made after the closing of the workshop.
How to get to Florence
By plane: The Amerigo Vespucci Airport is just 3 km from the center of Florence and offers service to Europe's main cities, as well as the Rome and Milan airports with connections to all international destinations. There is also a shuttle service with hourly departures (price Lit. 6.000, tickets can be purchased at the airport espresso bar or on board). Taxis are available on the "Arrivals" side of the airport, the average fair to downtown Florence is Lit. 30.000. The Pisa airport (80 km from Florence) is linked to Florence by trains leaving every hour.
By train: Florence has excellent rail connections to the major Italian and European cities via the Eurostar trains. The Santa Maria Novella Railroad Station is in the center of the city.
By car - Parking: You can access the city by car although the historic center is closed to private traffic. You can get to your hotel by showing the booking voucher, and then you can park in the hotel's nearby garages. The main underground public parking facilities (hourly or daily rates) are located under the Santa Maria Novella Railroad Station, under the Parterre in Piazza della Libert? and at the Fortezza da Basso.
Taxis: The minimum fare for a cab ride in Florence is Lit. 7.000. The radio cab numbers are: 055 4798; 055 4242 and 055 4390. There are cab stands at various strategic points throughout the city. You cannot hail cabs on the street.
Local Scientific Organizer Dr. Federico Magnani CNR - IMGPF Via A. Vannucci, 13 - 50134 Florence (Italy) Phone ++39 055 461453 - Fax ++39 055 486604 E mail: federico at imgpf.fi.cnr.it
Organizing Secretariat Enic Viale Amendola, 20 - 50121 Florence (Italy) Phone ++39 055 240275 - Fax ++39 055 2345078 E mail: chiara at egr.it- Website: http://www.egr.it
Prof. Satu Huttunen, Finland
Dr. Jurg Bucher, Switzerland
Prof. Bjorn Sundberg, Sweden
Prof. Paul G. Jarvis, United Kingdom
Prof. Rainer Matyssek, Germany
Prof. Giuseppe Scarascia Mugnozza , Italy
Prof. Marco Borghetti , Italy
Dr. Federico Magnani, Italy
CNR - National Research Council, Italy
SISEF - Italian Society of Silviculture and Forest Ecology
Italian Academy of Forest Science